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Moreover, studies on the ionic form of Cu absorbed by grass roots have yielded conflicting results, suggesting three not mutually exclusive models for Cu uptake (Figure 2): 1) Uptake of Cu + (Cu(I)) : Evidence from (Jouvin et al., Reference Jouvin, Weiss, Mason, Bravin, Louvat, Zhao, Ferec, Hinsinger and Benedetti2012) suggests that, as in non-graminaceous plants, Cu(II) is reduced to Cu(I) before uptake into the root epidermal cells of grasses