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The first who proposed prooxidant properties of carotenoids were Burton and Ingold (Burton and Ingold 1984)

Reduced autophagic activity not only leads to the delayed and abnormal accumulation of denatured proteins and dysfunctional organelles but also, through various pathways, results in endothelial dysfunction and intimal thickening, exacerbating vascular aging

The mechanism involves positive regulation of the expression of ALDHs and cytoprotective enzymes (e.g

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doi: 10.1016/j.bbrc.2022.01.106 201 MuriJ.KopfM

Additionally, GSH shifts the melanogenesis pathway away from eumelanin (the darker brown/black pigment) toward pheomelanin (the lighter red/orange pigment) by interacting with dopaquinone intermediates in the Raper-Mason pathway