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Catalyzed by glutathione synthetase ( GSS ), this step is generally not rate-limiting under normal conditions but GSS variants that reduce enzyme activity can become limiting when step 1 is driving high -GluCys throughput, or when glycine availability is restricted

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In the case of ischemic stroke, reactive astrocytes also contribute to BBB permeability through the Wnt/-catenin pathway as demonstrated in an Nhe1 conditional knockout mouse model 119

Hayashi, H., Sakai, K., Baba, H