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This metabolic inflexibility in Insr P1195L/ + mice could be also due to compensatory increase in fat utilization in fat and skeletal muscle during dark phase caused by impaired glucose uptake in these tissues

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The high Ca 2+ level in cytoplasm with mitochondria dysfunction and the low intracellular NAD + level both leads to the Warburg effect

(source) Valentino MA, Lin JE, Waldman SA