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50 mM Tris, 150 mM NaCl, pH 7.4) containing 0.5% (v/v) phosphatase inhibitor cocktail 2 (P5726, Sigma), 2% (w/v) protease inhibitors (Complete EDTA-free, 05056489001, Roche) and 5% (v/v) DNase (D5025, Sigma)

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3.2.1.21), an additional extraction was carried out using 50 mM Na-phosphate buffer (pH 5.0)